7 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 6 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 5 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 4 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 3 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 2 Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada. 1 Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada.13 Institute for the Oceans and Fisheries, University of British Columbia, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada. 12 Department of Botany, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada. 11 Department of Microbiology and Immunology, University of British Columbia, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada. 10 Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2207 Main Mall, Vancouver, BC V6T 1Z4, Canada. 9 Pacific Biological Station, Fisheries and Oceans Canada, 3190 Hammond Bay Rd, Nanaimo, BC V9T 6N7, Canada. 8 Biology Department, University of Victoria, 3800 Finnerty Rd, Victoria, BC V8P 5C2, Canada.